How will the sudden loss of light during Monday’s solar eclipse affect members of the animal kingdom?

While researchers already have some idea, they’ll nevertheless spend those four-and-a-half minutes of darkness not looking up, but observing the behaviors of creatures both great and small.

Zoos in Indiana, Arkansas, Ohio, and Texas all plan on monitoring their respective residents’ behaviors as the rare meteorological event unfolds.

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The solar eclipse will be visible from parts of New York — maybe even the city.

Because eclipses are such infrequent meteorological events, there isn’t a wealth of reliable data on the effects they have on birds, bugs, lizards, and animals that isn’t largely anecdotal.

But scientists did learn quite a bit about how different species respond to these rare occurrences during the “Great American Eclipse” of August 2017.

During that total solar eclipse, zoologists and animal behaviorists at the Riverbanks Zoo in Columbia, South Carolina, documented how 17 different species reacted to the phenomenon.

Their findings were published in the journal Animals, and suggest the sudden light shift creates a conflict within animals, even domesticated ones, between their internal rhythm and the world around them.

While 25% of the zoo’s creatures — including the Komodo dragon — exhibited signs of anxiety with the immediate onset of night, 75% began simply engaging in their nighttime routines.

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Female gorillas, for instance, “approached the entrance to their indoor enclosure in their typical hierarchical order from the far side of their exhibit,” the paper explains — “a behavior repeated every evening prior to their being secured inside for the night.”

Other creatures, like the typically-docile Galapagos tortoise, went on a mating frenzy and then “gazed up at the sky” as the light started returning.

For Monday’s eclipse, many of the same researchers will be holed up at the Houston Zoo, where they’re still looking for volunteers to observe animals during the eclipse. NASA is also seeking volunteers living within the path of totality and on its edge to help them document the effects of the eclipse on all forms of life.

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Nate Bickford, an animal researcher at Oregon Institute of Technology, told The Post the abrupt arrival of darkness had wild animals exhibiting the same sorts of behaviors they typically display when there’s a fast-approaching storm.

“So, a lot of the wildlife respond to the rapid darkening as though a powerful, nor’easter type storm coming through,” said Bickford, who studied animal behaviors in Nebraska in 2017.

Bickford said analyzing the different creature’s responses also provided insight into which ones rely on barometric pressure to gauge the arrival of oncoming storms, and which rely on the shift in light.

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“The ones that use barometric pressure, they don’t budge,” Bickford said with a laugh.

Expect songbirds to change course midflight, as they head back to their roosts, Bickford said. You may also spot spiders re-weaving their webs, or hear crickets start to chirp.

How might your pets react, and what, if anything, should you be doing to protect them during the eclipse? If your pet has an regular evening routine, Dr. Sun Kim of Cornell University’s Duffield Institute for Animal Behavior told The Post they may start acting like its bedtime.

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“Unless a pet doesn’t have situational anxiety, an owner may not need to be concerned about” the eclipse, Kim said. “For pets with a rare anxiety condition, such as nighttime anxiety, then giving situational anti-anxiety medications would be recommended.”

Kim said to keep them safe, keep your pets inside, and “give them fun things to do” while you go check out the eclipse. “Giving long-lasting food toys will be ideal for pets.”

Manhattan vet Dr. Lisa Lippman agrees that the safest bet for pet owners is to keep them inside.

“You don’t encourage them to look up or hold up treats,” Lippman said. “You don’t want to be taking animals with you to big outings with a lot of excited people.”

According to Bickford, there’s one species in particular that truly goes wild for eclipses.

“The organism that has the biggest response is human beings,” he said. “It’s fun to watch humans during the eclipse. They’re just excited, right? Yelling and screaming, in a positive way — and primates, they do something very similar to that.”